Interbull Routine Genetic Evaluation for Dairy Production Traits
The latest routine international evaluation for dairy
production traits took place as scheduled at the Interbull Centre. Data from
twenty five (25) countries were included in this evaluation. Results were
distributed on August 9, 2004, to designated representatives.
International genetic evaluations for milk, fat and protein yields of bulls from Australia, Austria, Belgium, Canada, Czech Republic, Denmark, Estonia, Finland, France, Germany, Hungary, Ireland, Israel, Italy, Japan, Netherlands, New Zealand, Norway, Poland, Rep. of South Africa, Slovenia, Spain, Sweden, Switzerland, the United Kingdom, and the United States of America were computed. Ayrshire, Brown Swiss, Guernsey, Holstein, Jersey and Simmental breed data were included in this evaluation.
New data (national genetic evaluations) included in this evaluation
Changes in previous test-run NOT introduced:
Changes in Interbull procedures
Data and method of analysis
Data were national genetic evaluations of AI sampled bulls
with daughters in at least 10 herds. Table 1 presents the amount of data
included in this Interbull evaluation. The difference between "No. Records" and
"No. Bulls" is explained by common bulls with proofs in more than one country.
The entry "Publishable Proofs" reflects bulls whose international evaluations
were distributed to service customers. The difference between "No. Bulls" and
"Publishable Proofs" is explained by bulls not meeting the minimum criterion for
official publication in the country of origin.
Table 2 and table 3 presents the current definition of the reference (genetic) base and date of evaluation, respectively, as supplied by each country in the dairy-production proof file.
Estimated genetic parameters are shown in APPENDIX I and the corresponding number of common bulls are listed in APPENDIX II. Parameters are listed by breed, trait and country.
National proofs were first de-regressed within country and then analysed jointly with a linear model including the effects of evaluation country, genetic group of bull and bull merit. Different breeds were analysed separately. Heritability estimates used in both the de-regression and international evaluation were as in each country's national evaluation (Table 4).
Ancestor-bulls without own proofs were traced back two generations from the oldest bulls with proofs in order to increase across country connections and account for the effect of selection.
Genetic groups were defined according to unidentified parents by national origin, breed and birth year of the bull and path of selection (sire, maternal grand-sire, maternal grand-dam). Birth year grouping was by 1-5 year periods. Small groups (generally consisting of less than 10 bull) were combined.
The international genetic evaluation procedure is based on international work described in the following scientific publications:
International genetic evaluation computation:
Schaeffer. 1994. J. Dairy Sci. 77:2671-2678
Klei, 1998. Interbull Bulletin 17:3-7
Verification and Genetic trend validation:
Klei et al., 2002. Interbull Bulletin 29:178-182.
Boichard et al., 1995. J. Dairy Sci. 78:431-437
Fikse and Banos, 2001. J. Dairy Sci. 84:1759-1767
De-regression and genetic parameter estimation
Sigurdsson and G. Banos. 1995. Acta Agric. Scand. 45:207-219
Jairath et al. 1998. J. Dairy Sci. Vol. 81:550-562
Genetic parameter estimation:
Sigurdsson et al., 1996. Acta Agric. Scand. 46:129-136
Sullivan, 1999. Interbull Bulletin 22:146-148
Jorjani et al., 2003. J. Dairy Sci. 86:677-679
Weigel and Banos. 1997. J. Dairy Sci. 80:3425-3430
International reliability estimation
Harris and Johnson. 1998. Interbull Bulletin 17:31-36
Genetic correlation estimation procedure
Estimation of genetic correlations among countries takes
place in test-runs only, when new or modified data are submitted from a country,
according to the following procedure (as per Interbull technical workshop of
April 1995, Uppsala, Sweden):
Several subsets of countries are analysed. At the most 10 countries at a time are included in each subset. Countries that are major link contributors (judged from the number and origin of common bulls with multiple national evaluations) are always included in these subsets. If multiple genetic correlation estimates are computed for a country pair, the highest estimate is kept, as per Sigurdsson et al (1996) showing that genetic correlations may be under-estimated but not over-estimated by the method used.
In some cases sufficient links between countries may be missing, resulting in close to zero genetic correlation estimates. If no reasonable correlations can be estimated via indirect links with third countries, one the the following procedures is followed:
a) Estimates from another breed for the country pair are used, if applicable
b) Product moment correlations of common bulls, adjusted for national evaluation accuracy are used (not a very frequent practice, since presence of common bulls will likely result in reasonable correlation estimates using the approximate REML method of Sigurdsson et al)
c) Estimates from the low end of the correlation distribution are assigned; these would normally range from .86 to .89 between two North Hemisphere countries and from .75 to .78 between a North and a South Hemisphere country; the model of national evaluation is also taken into consideration (countries with similar national evaluation models are assigned higher genetic correlation estimates)
Since genetic correlation estimates are not derived simultaneously, the full covariance matrix need to be bent in order to ensure it's positive definite.
Efforts to improve the procedure are currently under way. The use of covariance structure of models that include genetic and non-genetic (eg, national evaluation model, management practice etc) components in determining correlation estimates between weakly or non-linked countries/populations is being studied.
If a country is not linked to the other countries in the evaluation system, its data are not included in the international genetic evaluation.
Publication of Interbull evaluations
Results were distributed by the Interbull Centre to
designated representatives in each country. The international evaluation file
comprised international proofs expressed on the base and unit of each country
included in the analysis. Such records readily provide more information on bull
performance in various countries, thereby minimising the need to resort to
At the same time, all recipients of Interbull results are expected to honour the agreed code of practice, decided by the Interbull Steering Committee, and only publish international evaluations on their own country scale. Evaluations expressed on another country scale are confidential and may only be used internally for research and review purposes.
All recipients are also expected to follow the agreed guidelines for advertising genetic merit. The guidelines has been distributed to all members and is available on the Interbull homepage (http://www.interbull.org) under "Publications and Documentation/General information".
Next routine international evaluation
The next routine international evaluation for dairy production, udder health, conformation and longevity traits is scheduled for November 2004. New data for the November routine run should reach the Interbull Centre not later than Tuesday October 26, 2004, 17:00 Central European Time (CET); in any case, the most recently received data will be considered. The target time for distribution of results for internal review to designated representatives, who have signed a confidentiality agreement with the Centre, is Thursday November 4, 2004. Official release date of the results is targeted to be Monday November 8, 2004.
Next test international evaluation
The next test international evaluation for dairy
production, conformation and udder health traits is scheduled for September
2004. The next test evaluation for direct longevity for the Holstein breed is
March 2005. The first test evaluation for direct longevity (breeds other than
Holstein) and calving traits (Holstein only) is scheduled for September 2004.
Countries wishing to enter the system for the first time or planning to submit
new information (modified national evaluation procedure, new breeds etc) for
following routine evaluations must have their data tested in this test-run.
Deadline for sending data and validation results to the Interbull Centre for the September 2004 test evaluation is September 1, 2004 at 17:00 CET.
Means of result distribution from the Interbull Centre
Electronic exchange of data is probably more efficient
than anything else. Currently most countries in the service have established
internet connections and receive international evaluation results on the day of
their release. Subscribers to the service that do not already exercise this
option are encouraged to consider it and establish an internet connection and
ftp account. When such accounts are available, please provide their
specifications to the Interbull Centre. Until then, data will be delivered on
For more information about the international genetic evaluation service please contact the Interbull Centre: address: SLU Box 7023, 750 07 Uppsala, Sweden; fax: +46-18-672648;