INTERBULL Routine Genetic Evaluation for Dairy Production Traits

February 2000

Introduction

The latest routine international evaluation for dairy production traits took place as scheduled at the Interbull Centre. Data from twenty five (25) countries were included in this evaluation. Results were distributed on February 14, 2000, to designated representatives in the twenty five countries currently subscribing to the service.

International genetic evaluations for milk, fat and protein yields of bulls from Australia, Austria, Belgium, Canada, Czech Republic, Denmark, Estonia, Finland, France, Germany, Hungary, Ireland, Israel, Italy, Netherlands, New Zealand, Norway, Poland, Rep. of South Africa, Slovenia, Spain, Sweden, Switzerland, the United Kingdom, and the United States of America were computed. Ayrshire, Brown Swiss, Guernsey, Holstein, Jersey and Simmental breed data from 8, 10, 4, 26, 7 and 8 bull populations, respectively, were included in this evaluation.

New data (national genetic evaluations) included in this evaluation:

• New national genetic evaluations for Holstein and Ayrshire were computed in Sweden using an animal model (previously sire/MGS model) and included in this Interbull evaluation.
• The most recent data from 3 countries in the Holstein evaluation and from 1 country in the Ayrshire evaluation had to be excluded due to increases in national genetic evaluation variances leading to inflated MACE parameters with estimated sire variances severely exceeding the range set in the Interbull code of practice (with amendments). Such increases indicate changes in national evaluation models, affecting national evaluation results, that need to be tested in a test-run.

  Data from November 1999 was used in these cases.

  Countries are reminded that a set of programs has been made available, on the Interbull ftp-server, that can be used as a guidance to assess if (minor) changes in national evaluation models require a test-run.

• A new, unannounced, identification system for Black & White Holstein animals was introduced in Switzerland. Unfortunately, mistakes were made at the introduction that went unnoticed in the screening process at the Interbull Centre, and some foreign pedigree information was affected. About 17% of HOLCHE bulls had erroneous foreign sire id's and about 40% had erroneous MGS id's. Thus, bulls with wrong sire or MGS id's will not contribute to their relatives breeding values abroad, and vice versa.

  Note that the Swiss Red Holstein population was not affected.

Changes in the INTERBULL evaluation:

• Minimum birth year of bulls included in the international evaluation for dairy production and conformation traits was updated by 1 year, a procedure that was introduced at the February routine evaluation in 1998. The objective is to maintain a constant window of recent data across country on which international evaluations are based, in light of theoretical evidence that such recent data best describe the current situation in changing populations and more accurately assess the genetic merit of young bulls. Therefore, in this evaluation run minimum birth year was set to 1983 in the Holstein evaluation and 1978 in the evaluation of other breeds. All identifiable pedigree information was included in each evaluation.

Data and method of analysis

Data were national genetic evaluations of AI sampled bulls with daughters in at least 10 herds. Table 1 presents the type and amount of data included in this Interbull evaluation. The difference between "No. Records" and "No. Bulls" is explained by common bulls with proofs in more than one country. The entry "Publishable Proofs" reflects bulls whose international evaluations were distributed to service customers. The difference between "No. Bulls" and "Publishable Proofs" is explained by bulls not meeting the minimum criterion for official publication in the country of origin.

Table 2 presents the current definition of the reference (genetic) base as supplied by each country in the dairy-production proof file.

Estimated genetic parameters are shown in APPENDIX I. Parameters are listed by breed, trait, and country.

National proofs were first de-regressed within country and then analysed jointly with a linear model including the effects of evaluation country, genetic group of bull and bull merit. Different breeds were analysed separately. Heritability estimates used in both the de-regression and international evaluation were as in each country's national evaluation (Table 1).

Ancestor-bulls without own proofs were traced back two generations from the oldest bulls with proofs in order to increase across country connections and account for the effect of selection.

Genetic groups were defined according to unidentified parents by national origin, breed and birth year of the bull and path of selection (sire, maternal grand-sire, maternal grand-dam). Birth year grouping was by 1-5 year periods. Small groups (generally consisting of less than 10 bull) were combined.

Scientific literature

The international genetic evaluation procedure is based on international work described in the following scientific publications:

1. International genetic evaluation computation

L. Schaeffer. 1994. J. Dairy Sci. 77:2671-2678

2. Genetic trend validation

D. Boichard et al. 1995 J. Dairy Sci. 78:431-437

3. De-regression and genetic parameter estimation

A. Sigurdsson and G. Banos. 1995. Acta Agric. Scand. 45:207-219

A. Sigurdsson et al. 1996. Acta Agric. Scand. 46:129-136

P. Rozzi and L. Schaeffer. 1996. Interbull Technical Workshop, Verden, Germany, 25-26 Nov

4. Time edits

K. Weigel and G. Banos. 1997. J. Dairy Sci. 80:3425-3430

5. International reliability estimation

B. Harris and D. Johnson. 1998. Interbull Bulletin 17:31-36

Genetic correlation estimation procedure

Estimation of genetic correlations among countries takes place in test-runs only, when new or modified data are submitted from a country, according to the following procedure (as per Interbull technical workshop of April 1995, Uppsala, Sweden):

Step 1:

• Several subsets of countries are analysed. At the most 10 countries at a time are included in each subset. Countries that are major link contributors (judged from the   number and origin of common bulls with multiple national evaluations) are always included in these subsets. If multiple genetic correlation estimates are computed for a country pair, the highest estimate is kept, as per Sigurdsson et al (1996) showing that genetic correlations may be under-estimated but not over-estimated by the method used.
  

Step 2:

• In some cases sufficient links between countries may be missing, resulting in close to zero genetic correlation estimates. If no reasonable correlations can be estimated via indirect links with third countries, one of the following procedures is followed:
• a) Estimates from another breed for the country pair are used, if applicable
• b) Product moment correlations of common bulls, adjusted for national evaluation accuracy are used (not a very frequent practice, since presence of common bulls will likely result in reasonable correlation estimates using the approximate REML method of Sigurdsson et al)
• c) Estimates from the low end of the correlation distribution are assigned; these would normally range from .86 to .89 between two North Hemisphere countries and from .75 to .78 between a North and a South Hemisphere country; the model of national evaluation is also taken into consideration (countries with similar national evaluation models are assigned higher genetic correlation estimates)
  

Step 3:

• Since genetic correlation estimates are not derived simultaneously, the full covariance matrix need to be bent order to ensure it's positive definite; bending applies to estimates from Step 2 only.
  

APPENDIX I indicates estimates from Step 2c of the procedure.

Efforts to improve the procedure are currently under way. The use of covariance structure of models that include genetic and non-genetic (eg, national evaluation model, management practice etc) components in determining correlation estimates between weakly or non-linked countries/populations is being studied.

If a country is not linked to the other countries in the evaluation system, its data are not included in the international genetic evaluation.

Publication of INTERBULL evaluations

Results were distributed by the Interbull Centre to designated representatives in each country. The international evaluation file comprised international proofs expressed on the base and unit of each country included in the analysis. Such records readily provide more information on bull performance in various countries, thereby minimising the need to resort to conversions.

At the same time, all recipients of Interbull results are expected to honour the agreed code of practice, decided by the Interbull Steering Committee, and only publish international evaluations on their own country scale. Evaluations expressed on another country scale are confidential and may only be used internally for research and review purposes.

Use of INTERBULL evaluations in various countries

Updated information on national policies regarding use of Interbull evaluations in countries participating with data and release dates of national and Interbull evaluation results in various countries, is pending on results of an ongoing survey.

INTERBULL conversions

Since all data are now becoming available, conversions are not as useful as before. When an international proof is available, customers of the service are encouraged to use this instead of a converted proof. Conversion coefficients were, however, computed from international evaluation results and may be considered "Estimated Interbull Conversions" for the interim until the next international evaluation run.

Next routine international evaluation

The next routine international evaluation for dairy production and conformation traits is scheduled for May 2000. New data for that run should reach the Interbull Centre not later than April 28, 2000, 17:00 Central European Time (CET); in any case, the most recently received data will be considered. Results will be distributed on May 8, 2000.

Next test international evaluation

The next test international evaluation for dairy production and conformation traits is scheduled for March 2000. Countries wishing to enter the system for the first time or planning to submit new information (modified national evaluation procedure, new breeds etc) for the following routine evaluation must have their data tested in this test-run.

Deadline for sending new data to the Interbull Centre for the next test-run is March 1, 2000.

Means of result distribution from the Interbull Centre

Electronic exchange of data is probably more efficient than anything else. Currently most countries in the service have established internet connections and receive international evaluation results on the day of their release. Subscribers to the service that do not already exercise this option are encouraged to consider it and establish an internet connection and ftp account. When such accounts are available, please provide their specifications to the Interbull Centre. Until then, data will be delivered on CD-ROM's.

For more information about the international genetic evaluation service please contact the INTERBULL Centre: address: SLU Box 7023, 750 07 Uppsala, Sweden;  fax: +46-18-672648;

e-mail: Ulf.Emanuelson_at_hgen.slu.se

e-mail: Georgios.Banos_at_hgen.slu.se

e-mail: Freddy.Fikse_at_hgen.slu.se

e-mail: Hossein.Jorjani_at_hgen.slu.se

e-mail: Jan.Philipsson_at_hgen.slu.se

e-mail: Dan.Englund_at_hgen.slu.se